Joe Beck has lost the love of his life thanks to a Pixie"s Curse and he now needs to figure out how to get his girl back, learn more about the mysterious Pixies and undo a bad deed he did long ago.

Fievel is a young Russian mouse separated from his parents on the way to America, a land they think is without cats. When he arrives alone in the New World, he keeps up hope, searching for his family, making new friends, and running and dodging the cats he thought he"d be rid off.